Course # CZMT-0671 CRN: 26090
* For Undergraduate and general interest credit only
Course # OCUG-3400* CRN: 26099
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A pair of Blackfin Reef Sharks (Carcharhinus melanopterus) cruises among the pilings of a resort dock at North Male' Atoll, Maldives. Humans and sharks come into contact with ever-increasing frequency; sharks almost always fare worse as a consequence of encounters with us. |
Course Description:
Sharks and rays (collectively termed "elasmobranchs") are creatures of biological elegance and perennial fascination. In recent years, new technologies have revealed fascinating details about the heretofore secret lives of elasmobranchs as well as demonstrated their population declines on an unprecedented and global scale.This unique 12-week course is a comprehensive, interdisciplinary introduction to the evolution, biology, ecology, and conservation of elasmobranch fishes.
Topics covered include evolution, classification, diversity, anatomy, physiology, sensory biology, locomotion, feeding, reproduction, life history, behavior, distribution, ecology, and conservation of sharks and rays.
Student Base:A credit course for graduate students, advanced undergraduate students (transfer credit is available for students from other institutions) and general interest distance learners. Elasmobranchs are important components in virtually all marine ecosystems and in a few tropical freshwater ones as well. Sharks and rays are commercially exploited and vulnerable to directed fisheries, bycatch, and anthropogenic habitat erosion. This course will play an important role in enriching the educational background and professional understanding of marine biology, zoology, or coastal management degree students.
Credits: 3
Prerequisites: If taken toward an undergraduate degree, it is recommended that students have completed at least second-year biology courses. No prerequisites for those taking the course for general interest.
![]() A Blotched Fantail Ray (Taeniura meyeni) with an entourage of trevally (Carangidae) at North Male' Atoll, Maldives |
This course 1) integrates and amplifies scientific learning from basic biology, physics, and chemistry, and 2) provides students with a comprehensive, well-rounded, and up-to-date understanding of the biology of elasmobranch fishes, 3) building toward pursuit of careers in marine ecology, zoology, ichthyology, and fisheries management.
Class Format:
This course is taught on-line, based on peer-reviewed literature in the anthology selected as the course textbook (see below). Selected readings will be assigned each week for background reading, and classroom discussions. The estimated time commitment is five to ten hours per unit, on average, to complete readings and participate in course activities.
Course Materials:
Course content is delivered as self-guided narrated and illustrated lectures. For undergraduate students and general interest learners, these are augmented by the text, Sharks in Question by Victor Springer and Joy Gold (Smithsonian Institution Press, 1989); for graduate students, course content is amplified and extended by the text, Biology of Sharks and Their Relatives edited by Carrier, Musick, and Heithaus (CRC Press, 2004). Additional supplementary material includes on-line video clips and use of selected Internet resources. Interaction with the course instructor is via E-mail and participation in on- line discussions centering on topics set by the instructor.
Course Resources:
Print and Web-CT.
Unit Plan:
| Week | Topic | Week | Topic |
| 1 | Evolution | 7 | Feeding |
| 2 | Classification | 8 | Reproduction |
| 3 | Diversity | 9 | Age and Growth |
| 4 | Form and Function | 10 | Brain and Behavior |
| 5 | Sensory Biology | 11 | Distribution and Ecology |
| 6 | Swimming and Body Form | 12 | Conservation |
Attendance:
Students are expected read all material provided. Attendance will be gauged by participation in on-line discussion and submission of research projects.
![]() A Great White Shark (Carcharodon carcharias) off the Western Cape, South Africa |
Fieldtrips:
If enough participation can be secured, a formal shark-watching field trip in the Florida-Bahamas region can be arranged, in
which students will apply what they have learned about shark behavior in the course.
Assessment:
For undergraduate students, participation in on-line discussions (20%), an on-
line mid-term test (short answer, 25%), a term project on an elasmobranch-
related topic of the student's choice but subject to approval by the course
instructor (25%), and a final examination consisting of short answers plus a
choice of two of six integrative essay questions, to be submitted via E-mail
attachment and graded by the instructor (30%).
For graduate students, participation in on-line discussions (20%), a term
project on an elasmobranch-related topic of the student's choice but subject to
approval by the course instructor (30%), and a final examination consisting of
three integrative essay questions set by the instructor, to be submitted via E-
mail attachment and graded by the instructor (50%).
| UNIT | TOPICS | UNIT | TOPICS |
| 1 |
Evolution What are Elasmobranchs? Why Study Elasmobranchs? Evolutionary Processes Fossilization Geologic Time Interpreting Fossils The Earliest Sharks Carboniferous Sharks Hybodonts Neoselachians |
7 |
Feeding Foraging Strategy Predatory Cycle Dietary Categories Predatory Strategies Prey Handling Cooperative Hunting Filter-Feeding Facultative Ectoparasitism Scavenging Food Requirements Trophic Level |
| 2 |
Classification Why Classify Organisms? Systematics Sources of Data Development of Elasmobranch Classification Modern Elasmobranch Classification |
8 |
Reproduction Reproductive Strategy Reproductive Structures Male Reproductive Anatomy Female Reproductive Anatomy Reproductive Behavior Hormonal Regulation Pheromones Courtship Rituals Mate Selection Cooperative Mating Reproductive Modes Development Nursery Areas Fecundity |
| 3 |
Diversity Phyletic Diversity Survey of Galeomorph Elasmobranchs Survey of Squalomorph Elasmobranchs Survey of Batoid Elasmobranchs Ecological Diversity Habitat Types Ecomorphotypes |
9 |
Age and Growth Age Determination Growth Stages Growth Rates Maximum Size Longevity |
| 4 |
Form and Function Squamation Pigmentation Teeth Endoskeleton Muscular System Respiratory System Circulation Digestion Osmoregulation Immunology Metabolism |
10 |
Brain and Behavior Central Nervous System Brain Organization Brain Weight vs. Body Weight Spinal Chord Activity Patterns Home Range Circadian Rhythms Seasonal Movements Migration Social Behavior Competition Social Organization of Populations Learning Intelligence |
| 5 |
Sensory Biology Mechanoreception Chemoreception Photoreception Electroreception Proprioception Sensory Integration |
11 |
Distribution and Ecology Oceanic Provinces Oceanic Zones Temperature Regimes Habitat Types Habitat Selection Distribution Patterns Zoogeography Ecology The Niche Concept Ecological Separation Ecological Role Population Size Symbionts Sources of Mortality Natural Anthropogenic |
| 6 |
Swimming and Body Form Hydrodynamics Drag Bouyancy Role of Fins Propulsion Styles Body Forms |
12 |
Conservation Falling Populations Lack of Information Demography Genetics Management Models Lack of Funding Systematics Life History Conservation Status IUCN Criteria Public Awareness Why Conserve Elasmobranchs? |
Grading:
All exercises, including exams and submissions must be E-mailed to instructor. Grading will take into consideration: content, organization, accuracy and quality of presentation. Late assignments will incur in a 20-point reduction (out of a possible maximum of 100).
About the Instructor:
The "Biology of Sharks and Rays" course was designed and taught by R. Aidan Martin, Director of the ReefQuest Centre for Shark Research (www.elasmo-research.org), a Research Associate of the Zoology Department of the University of British Columbia, and an Adjunct Professor of the Oceanographic Center of Nova Southeastern University. Aidan died unexpectedly on February 13, 2007 and the next offering of the course is on hold, pending discussions with his wife, Anne and Dr. Richard Spieler, Director of Academic Programs. For a memorial to Aidan, Click here.
If you register for Biology of Sharks and Rays, you will be expected to make contact with the co-ordinator by e-mail.
Please address requests for additional information regarding the Biology of Sharks and Rays distance education course to
Email: "Biology of Sharks and Rays" <de-sharks_1 barney mail fred ocean fred nova fred edu>