NSUOC Home

Biology of Sharks and Rays

Course # CZMT-0671 CRN: 26090
Course # OCUG-3400* CRN: 26099

* For Undergraduate and general interest credit only

Click here for information regarding applying for admission for distance education courses

 


A pair of Blackfin Reef Sharks (Carcharhinus melanopterus) cruises among the pilings of a resort dock at North Male' Atoll, Maldives. Humans and sharks come into contact with ever-increasing frequency; sharks almost always fare worse as a consequence of encounters with us.

Course Description:

Sharks and rays (collectively termed "elasmobranchs") are creatures of biological elegance and perennial fascination. In recent years, new technologies have revealed fascinating details about the heretofore secret lives of elasmobranchs as well as demonstrated their population declines on an unprecedented and global scale.This unique 12-week course is a comprehensive, interdisciplinary introduction to the evolution, biology, ecology, and conservation of elasmobranch fishes.

Topics covered include evolution, classification, diversity, anatomy, physiology, sensory biology, locomotion, feeding, reproduction, life history, behavior, distribution, ecology, and conservation of sharks and rays.

Student Base:

A credit course for graduate students, advanced undergraduate students (transfer credit is available for students from other institutions) and general interest distance learners. Elasmobranchs are important components in virtually all marine ecosystems and in a few tropical freshwater ones as well. Sharks and rays are commercially exploited and vulnerable to directed fisheries, bycatch, and anthropogenic habitat erosion. This course will play an important role in enriching the educational background and professional understanding of marine biology, zoology, or coastal management degree students.

Credits: 3

Prerequisites: If taken toward an undergraduate degree, it is recommended that students have completed at least second-year biology courses. No prerequisites for those taking the course for general interest.


A Blotched Fantail Ray (Taeniura meyeni) with an entourage of trevally
(Carangidae) at North Male' Atoll, Maldives
 
Course Objectives:

This course 1) integrates and amplifies scientific learning from basic biology, physics, and chemistry, and 2) provides students with a comprehensive, well-rounded, and up-to-date understanding of the biology of elasmobranch fishes, 3) building toward pursuit of careers in marine ecology, zoology, ichthyology, and fisheries management.

Class Format:

This course is taught on-line, based on peer-reviewed literature in the anthology selected as the course textbook (see below). Selected readings will be assigned each week for background reading, and classroom discussions. The estimated time commitment is five to ten hours per unit, on average, to complete readings and participate in course activities.

Course Materials:

Course content is delivered as self-guided narrated and illustrated lectures. For undergraduate students and general interest learners, these are augmented by the text, Sharks in Question by Victor Springer and Joy Gold (Smithsonian Institution Press, 1989); for graduate students, course content is amplified and extended by the text, Biology of Sharks and Their Relatives edited by Carrier, Musick, and Heithaus (CRC Press, 2004). Additional supplementary material includes on-line video clips and use of selected Internet resources. Interaction with the course instructor is via E-mail and participation in on- line discussions centering on topics set by the instructor.

Course Resources:

Print and Web-CT.

Unit Plan:
 
Week Topic Week Topic
1 Evolution 7 Feeding
2 Classification 8 Reproduction
3 Diversity 9 Age and Growth
4 Form and Function 10 Brain and Behavior
5 Sensory Biology 11 Distribution and Ecology
6 Swimming and Body Form 12 Conservation

Attendance:

Students are expected read all material provided. Attendance will be gauged by participation in on-line discussion and submission of research projects.


A Great White Shark (Carcharodon carcharias) off the
Western Cape, South Africa
 

Fieldtrips:

If enough participation can be secured, a formal shark-watching field trip in the Florida-Bahamas region can be arranged, in which students will apply what they have learned about shark behavior in the course.

Assessment:

For undergraduate students, participation in on-line discussions (20%), an on- line mid-term test (short answer, 25%), a term project on an elasmobranch- related topic of the student's choice but subject to approval by the course instructor (25%), and a final examination consisting of short answers plus a choice of two of six integrative essay questions, to be submitted via E-mail attachment and graded by the instructor (30%).

For graduate students, participation in on-line discussions (20%), a term project on an elasmobranch-related topic of the student's choice but subject to approval by the course instructor (30%), and a final examination consisting of three integrative essay questions set by the instructor, to be submitted via E- mail attachment and graded by the instructor (50%).

Course Syllabus

UNIT TOPICS UNIT TOPICS
1 Evolution
What are Elasmobranchs?
Why Study Elasmobranchs?
Evolutionary Processes
Fossilization
Geologic Time
Interpreting Fossils
The Earliest Sharks
Carboniferous Sharks
Hybodonts
Neoselachians
7 Feeding
Foraging Strategy
Predatory Cycle
Dietary Categories
Predatory Strategies
Prey Handling
Cooperative Hunting
Filter-Feeding
Facultative Ectoparasitism
Scavenging
Food Requirements
Trophic Level
2 Classification
Why Classify Organisms?
Systematics
Sources of Data
Development of Elasmobranch Classification
Modern Elasmobranch Classification
8 Reproduction
Reproductive Strategy
Reproductive Structures
    Male Reproductive Anatomy
    Female Reproductive Anatomy
Reproductive Behavior
Hormonal Regulation
Pheromones
Courtship Rituals
Mate Selection
Cooperative Mating
Reproductive Modes
Development
Nursery Areas
Fecundity
3 Diversity
Phyletic Diversity
    Survey of Galeomorph Elasmobranchs
    Survey of Squalomorph Elasmobranchs
    Survey of Batoid Elasmobranchs
Ecological Diversity
    Habitat Types
    Ecomorphotypes
9 Age and Growth
Age Determination
Growth Stages
Growth Rates
Maximum Size
Longevity
4 Form and Function
Squamation
Pigmentation
Teeth
Endoskeleton
Muscular System
Respiratory System
Circulation
Digestion
Osmoregulation
Immunology
Metabolism
10 Brain and Behavior
Central Nervous System
Brain Organization
Brain Weight vs. Body Weight
Spinal Chord
Activity Patterns
Home Range
Circadian Rhythms
Seasonal Movements
Migration
Social Behavior
Competition
Social Organization of Populations
Learning
Intelligence
5 Sensory Biology
Mechanoreception
Chemoreception
Photoreception
Electroreception
Proprioception
Sensory Integration
11 Distribution and Ecology
Oceanic Provinces
Oceanic Zones
Temperature Regimes
Habitat Types
Habitat Selection
Distribution Patterns
Zoogeography
Ecology
The Niche Concept
Ecological Separation
Ecological Role
Population Size
Symbionts
Sources of Mortality
    Natural
    Anthropogenic
6 Swimming and Body Form
Hydrodynamics
Drag
Bouyancy
Role of Fins
Propulsion Styles
Body Forms
12 Conservation
Falling Populations
Lack of Information
Demography
Genetics
Management Models
Lack of Funding
Systematics
Life History
Conservation Status
IUCN Criteria
Public Awareness
Why Conserve Elasmobranchs?

Grading:

All exercises, including exams and submissions must be E-mailed to instructor. Grading will take into consideration: content, organization, accuracy and quality of presentation. Late assignments will incur in a 20-point reduction (out of a possible maximum of 100).

About the Instructor:

The "Biology of Sharks and Rays" course was designed and taught by R. Aidan Martin, Director of the ReefQuest Centre for Shark Research (www.elasmo-research.org), a Research Associate of the Zoology Department of the University of British Columbia, and an Adjunct Professor of the Oceanographic Center of Nova Southeastern University. Aidan died unexpectedly on February 13, 2007 and the next offering of the course is on hold, pending discussions with his wife, Anne and Dr. Richard Spieler, Director of Academic Programs. For a memorial to Aidan, Click here.

If you register for Biology of Sharks and Rays, you will be expected to make contact with the co-ordinator by e-mail. Please address requests for additional information regarding the Biology of Sharks and Rays distance education course to Email: "Biology of Sharks and Rays" <de-sharks_1 barney mail fred ocean fred nova fred edu>